Six stomatal types are recognized in the
Amaranthaceae viz.,
1.
anomocytic,
2.
anisocytic,
3.
diacytic,
4.
paracytic,
5.
hemiparacytic and
6.
brachyparacytic,
the last two types being in very low
frequencies(Padmini and Shanmukha Rao 1995).
The original terms of describing the
stomatal types are after Metcalfe & Chalk(1950),van
Cotthem(1970) and Dilcher(1974).
Anomocytic stomata: Monocyclic,subsidiaries
4 or more, indistinct from other epidermal cells,subsdisries
variable in position.
Anisocytic stomata: Monocyclic,
subsidiaties 3, distinct or indistinct from other epidermal
cells,one being smaller than the other two and variable in
position.
Diacytic stomata : Subsidiaries 2, with
conjoint walls at right angles or approximately right angles to
the guard cell,monocyclic, or occasionally one and a half
cyclic.
Paracytic stomata: Subsidiaries 2, stomata
completely surrounded by two parallel subsidiary cells with
conjoint walls towards the poles or approximately so, monocyclic
or occasionally one and a half cyclic.
Hemiparacytic stomata:
Monocyclic,subsidiaries 4-5, one distinct subsidiary parallel to
the long axis of the pore,longer or shorter than the guard
cells.
Brachyparacytic stomata:
Monocyclic,subsidiaries 4-5.two subsidiaries distinct, parallel
to the stoma, smaller in size when compared to the other
subsidiaries, the poles exposed to indistinct subsidiaries.
Stomatogenesis of 31 taxa
representing all the three tribes of Amaranthaceae were studied,
many of them for the first time.The ontogeny of anisocytic and
anomocytic is mesoperigenous dolabrat or unilabrate , whereas
the paracytic and diacytic follow exclusively mesogenous
dolabrate pattern, and the hemiparacytic and brachyparacytic
stomata develop through mesoperigenous dolabrate pattern.( if
this paragraph is given in the beginning it need not be
repeated)
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